But they face risks from Arctic development, a new study says.

Caribou and reindeer travel the longest distance while migrating on land, a new study shows. But they don’t move the most throughout the year; that distinction goes to gray wolves.

But all wide-ranging animals face dangers from human development, researchers say.

The study, published in Scientific Reports, sought to provide data for a long-held assertion, based on a study from 1989, that caribou travel the farthest distance. The researchers gathered data from GPS collars on terrestrial animals around the world, from caribou and wolves to camels and khulan (Mongolian wild ass).

They found that caribou migrate about 745 miles a year round-trip, clocking in at the longest terrestrial migration. (Whales and birds migrate much farther by sea and by air.)

But the researchers also found that for sheer distance traveled, gray wolves trek farther every year. One male wolf from Mongolia traveled 4,503 miles, the study found. Arctic foxes also outpaced caribou, traveling more than 3,600 miles.

These findings reveal that migration is a complicated concept. Read more at ArcticToday.